Ramphastos_tocoRamphastos_vitellinus_ariel_AmazoniaRamphastos_vitellinus_culminatusRamphastos_vitellinus_vitellinusRamphastos_vitellinus_ariel_southeastern_BrazilRamphastos_brevisRamphastos_dicolorusRamphastos_tucanus_cuvieriRamphastos_tucanus_tucanusRamphastos_swainsoniiRamphastos_ambiguusRamphastos_sulfuratusOsteologyMaximum ParsimonyBased on reanalysis of the original matrix.Weckstein_2005a