LebiasinaPiabucinaPyrrhulinaNannostomusCopellaHepsetusErythrinusHoplerythrinusHopliasCtenolucius_beaniCtenolucius_hujetaBoulengerella_lateristrigaBoulengerella_maculataBoulengerella_cuvieriBoulengerella_luciusBoulengerella_xyrekesOsteologyMaximum ParsimonyBased on reanalysis of the original matrix.Vari_1995a