Leopardus_pardalisPuma_concolorNeofelis_nebulosaPanthera_leoPanthera_oncaPanthera_pardusPanthera_tigrisPanthera_unciaPanthera_blytheaePanthera_atroxPanthera_palaeosinensisPanthera_spelaeaOsteologyMaximum ParsimonyBased on reanalysis of the original matrix.Tseng_etal_2013a