Archaeopteryx_lithographicaProtopteryx_fengningensisPengornis_houiApsaravis_ukhaanaGansus_yumenensisEnaliornisPasquiaornisChupkaornis_keraorumBrodavisBaptornis_advenusFumicollis_hoffmaniParahesperornis_alexiHesperornis_regalisHesperornis_gracilisIchthyornis_disparLithornisVegavis_iaaiCrypturellus_undulatusChauna_torquataAnas_platyrhynchosGallus_gallusPhasianus_colchicusOsteologyMaximum ParsimonyBased on reanalysis of the original matrix.Tanaka_etal_2018b