HyracotheriumPachynolophusDanjiangiaLambdotheriumEotitanopscf_EotitanopsPalaeosyopsBunobrontops_savageiMesatirhinus_juniusDesmatotitan_tukhumAcrotitan_ulanshirehDolichorhinus_hyogSphenocoelus_uintenMicrotitan_mongolFossendorhinus_diploMetarhinus_fluviatilisMetarhinus_abbottiMetarhinus_paterSthenodectes_incisivumTelmatherium_validusWickia_brevirhinusMetatelmatherium_ultiQufutitan_zhouiEpimanteoceras_formosusProtitan_grangeriProtitan_minorProtitanotherium_emargRhinotitan_kaiseniRhinotitan_andrewsiNanotitanops_shangPollyosbornia_altidensPygmaetitan_panxianenGnathotitan_berkeyiAktautitan_hippopotBrachydiast_transylvMetatitan_primusMetatitan_relictusMetatitan_khaitshinusNasamplus_progressusProtembolotherium_efrEmbolotherium_andrewsiEmbolotherium_grangeriPachytitan_ajaxDiplacodon_elatusParabrontops_gobiensisEubrontotherium_clarnProtitanops_curryiNotiotitanops_mississDianotitan_lunanensisDuchesneodus_uintensisMegacerops_coloradensisMegacerops_kuwagataParvicornis_occidentOsteologyMaximum ParsimonyBased on reanalysis of the original matrix.Mihlbachler_et_Demere_2010a