M_giganteusMetacryphaeus_tuberculatusPlesiomalvinella_bouleiPlesiomalvinella_pujraviiParabouleiaBouleiaaustralisbuddeaehaugidonegalenusgemellusgoeldiasperalacunaferaOsteologyMaximum ParsimonyBased on reanalysis of the original matrix.Lieberman_etal_1991a