ancestorCycloderma_aubryiPelochelys_bibroniAmyda_cartilagineusCyclanorbis_elegansRafetus_euphraticusApalone_feroxNilsonia_formosusCycloderma_frenatumAspideretes_gangeticusAspideretes_hurumPelochelys_indicaApalone_muticusLissemys_punctataCyclanorbis_senegalensisPelodiscus_sinensisApalone_spiniferusPalea_steindachneriDogania_subplanusTrionyx_triunguisPlastomenus_aff_thomasiiOsteologyMaximum ParsimonyBased on reanalysis of the original matrix.Joyce_et_Lyson_2010b