LeptictidaeOrycteropus_aferBos_taurusOvisOdocoileus_virginianusTragulusTursiopsDelphinapterus_leucasPhyseter_macrocephalusBalaenopteraHippopotamus_amphibiusChoeropsis_liberiensisSus_scrofaTayassu_tajacuCamelusLamaEquus_caballusCanisRattus_norvegicusVulpavusArctocyonHyopsodusPhenacodusMeniscotheriumHeptodonHyracotheriumMesohippusDiacodexis_pakistanensisDiacodexis_metsiacusHomacodon_vagansRaoellidaeBunomeryx_montanusGobiohyus_orientalisEntelodonElomeryx_armatusAmphirhagatherium_weigeltiPerchoerusAgriochoerusMerycoidodonMixtotheriumCebochoerusCainotheriumPoebrotheriumXiphodonEotylopus_reediLeptoreodon_marshiHeteromeryx_disparProtoceras_celerLeptomeryxHypertragulusAmphimeryxEoconodonAndrewsarchusHapalodectes_hetangensisHapalodectes_leptognathusDissacus_praenuntiusDissacus_navajoviusMongolian_DissacusAnkalagon_saurognathusSinonyx_jiashanensisPachyaena_giganteaPachyaena_ossifragaMesonyx_obtusidensSynoplotherium_caniusHarpagolestesPakicetusAmbulocetus_natansRemingtonocetus_harudiensisArtiocetus_clavisRodhocetusGeorgiacetus_vogtlensisProtocetus_atavusBasilosaurusOsteologyMaximum ParsimonyBased on reanalysis of the original matrix.Geisler_et_Uhen_2005a