Proganochelys_quenstedtiRhinochelys_pulchricepsSantanachelys_gaffneyiBouliachelys_suteriNotochelone_costataOcepechelon_bouyaiDesmatochelys_lowiiDesmatochelys_padillaiProtostega_gigasArchelon_ischyrosDermochelys_coriaceaEosphargis_breineriLepidochelys_olivaceaCaretta_carettaChelonia_mydasEretmochelys_imbricataLepidochelys_kempiiNatator_depressusPuppigerus_camperiArgillochelys_cuneicepsAllopleuron_hoffmanniToxochelys_spEochelone_brabanticaSolnhofia_parsoniJurassichelon_oleronensisPortlandemys_mcdowelliPlesiochelys_planicepsPlesiochelys_etalloniSandownia_harrisiBrachyopsemys_tingitanaLeyvachelys_cipadiChelydra_serpentinaMacrochelys_temminckiSternotherusStaurotypusKinosternon_suburum_hippocrepisDermatemys_mawiiBaptemys_wyomingensisEmarginachelys_cretaceaChrysemys_pictaEmys_orbicularisGeoclemys_hamiltoniiTestudoGopherus_polyphemusChelonoidis_spPlatysternon_megacephalumApalone_spiniferaPelodiscus_sinensisLissemys_punctataCarettochelys_insculptaAllaeochelys_libycaChelus_fimbriatusPhrynops_geoffranusElseya_dentataChelodinaAraripemys_barretoiPodocnemisPelomedusa_subrufaOrdosemys_sp_IVPP_V12092Judithemys_sukhanoviAnnemys_levensisAnnemys_latiensAnnemys_sp_IVPP_V18106Xinjiangchelys_wusuXinjiangchelys_radiplicatoidesDracochelys_bicuspisKirgizemys_hoburensisKirgizemys_dmitrieviSinemys_gameraSinemys_lensMeiolania_planicepsChubutemys_copelloiEubaena_cephalicaArundelemys_dardeniPleurosternon_bullockiiGlyptops_plicatulusKallokibotion_bajazidiEileanchelys_waldmanniKayentachelys_aprixAustralochelys_africanusOsteologyMaximum ParsimonyBased on reanalysis of the original matrix.Evers_et_Benson_2019a