Hypothetical_ancestorHyrachyus_eximiusHeptodonHelaletes_nanusDesmatotherium_intermediusDilophodon_minusculusColodon_occidentalisColodon_inceptusPlesiocolopirusProtapirusIrdinolophus_mongoliensisIrdinolophus_tuiensisTeleolophusRhodopagus_minimusTriplopus_cubitalisOsteologyMaximum ParsimonyBased on reanalysis of the original matrix.Dashzeveg_et_Hooker_1997a