ManouriaGopherus_agassiziiGopherus_berlandieriGopherus_brevisternusGopherus_canyonensisGopherus_edaeGopherus_flavomarginatusGopherus_hexagonatusGopherus_laticuneusGopherus_mohavetusGopherus_pansusGopherus_polyphemusGopherus_praecedensGopherus_vagusHesperotestudoStylemysTestudininaeOsteologyMaximum ParsimonyBased on reanalysis of the original matrix.Crumly_1993a