ArchaeopteryxJeholornisSapeornisConfuciusornisChangchengornisEoconfuciusornisProtopteryxOrienantiusEopengornisParapengornisJunornisConcornisElsornisEoalulavisCathayornisEocathayornisEoenantiornisGobipteryxIberomesornisLongipteryxLongirostravisNeuquenornisPengornisRapaxavisShanweiniaoVescornisVoronaSchizoouraJianchangornisArchaeorhynchusLongicrusavisApsaravisHongshanornisYanornisPatagopteryxYixianornisGansus_yumenensisGansus_zheniIchthyornisHesperornisParahesperornisVegavisAnasGallusBellulornisFortunguavisQilianiashenqiornisSulcavisBohaiornisParabohaiornisLongusunguisZhouornisCUGB_P1202GretcheniaoOsteologyMaximum ParsimonyBased on reanalysis of the original matrix.Chiappe_etal_2019a