AncestorHapalopsDeseadognathusProschismotheriumPseudorthotheriumEucholoeopsSchismotheriumPrepotheriumAmphiocnusPronothrotheriumMegatheriumNothrotheriopsAcratocnusCholoepusBradypusOsteologyMaximum ParsimonyBased on reanalysis of the original matrix.Carlini_et_Scillato-Yane_2004a