RhamphorhynchusPterodactylus_kochiPteranodonSantanadactylus_araripensisSantanadactylus_brasiliensis_HOLOSantanadactylus_brasiliensis_PARASantanadactylus_priceiSantanadactylus_spixiOrnithodesmusOrnithocheirusBHI_1276QuetzalcoatlusAzhdarchoTitanopteryxPteranodon_oregonensisDoratorhynchusDsungaripterusNyctosaurus_gracilisNyctosaurus_lamegoiOsteologyMaximum ParsimonyBased on reanalysis of the original matrix.Bennett_1989a