DromaeosauridaeArchaeopteryxJeholornis_primaSapeornisConfuciusornis_sanctusChangchengornisEoconfuciusornisCathyaornis_yandicaConcornisElsornisEoalulavisEocathayornisEoenantiornisGobipteryx_N_valifanoviIberomesornisLongipteryxLongirostravisProtopteryxRapaxavisShanweiniaoShenqiornisEopengornisPengornisAvisaurus_archibaldiA_gloriaeKaiparowitsSoroavisaurusIntiornisBauxitornisEnantiophoenixHalimornisNeuquenornisMystiornisArchaeorhynchusApsaravisYanornisPatagopteryxGansus_yumenensisLongicrusavisIchthyornisHesperornisAnasGallusOsteologyMaximum ParsimonyBased on reanalysis of the original matrix.Atterholt_etal_2018a