(Dromaeosauridae,(Archaeopteryx,(Jeholornis,(((Chongmingia,IVPP_V24194),(Sapeornis,((Pengornis,Eopengornis,Protopteryx,(Monoenantiornis,((Fortunguavis,(Linyiornis,(Eoenantiornis,(Concornis,Eoalulavis,Cathayornis,Gobipteryx,Neuquenornis,Vescornis,Qiliania,Piscivorenantiornis,Pterygornis,Cruralispennia,IVPP_V25033,(Eocathayornis,((Boluochia,Longipteryx),(Shanweiniao,(Longirostravis,Rapaxavis)))),(Elsornis,Dunhuangia))))),(Sulcavis,Shenqiornis,(Longusunguis,(Zhouornis,(Bohaiornis,Parabohaiornis))))))),(Archaeorhynchus,(Jianchangornis,((Schizooura,Bellulornis),((Songlingornis,(Yanornis,(Piscivoravis,(Apsaravis,Yixianornis,Iteravis,Gansus,Ichthyornis,(Vorona,Patagopteryx),((Enaliornis,(Baptornis_advenus,(Baptornis_varneri,(Hesperornis,Parahesperornsi)))),(Vegavis,(Anas,Gallus))))))),(Hongshanornis,(Parahongshanornis,(Longicrusavis,(Archaeornithura,Tianyuornis))))))))))),(Eoconfuciusornis_zhengi,(Confuciusornis_sanctus,Changchengornis_hengdaoziensis,Confuciusornis_dui,IVPP_V18929))))));