(Dimorphodon_macronyx,(Eudimorphodon_ranzii,(Anhanguera_piscator,Anhanguera_santanae,Anurognathus_ammoni,Arthurdactylus_conandoylei,Barbosania_gracilirostris,Campylognathoides_liasicus,Darwinopterus_linglongtaensis,DFMMh_slash_FV_500_dsungaripterid,Gegepterus_changae,Germanodactylus_rhamphastinus,Rhamphorhynchus_muensteri,Pterodactylus_antiquus,Nyctosaurus_gracilis,Pteranodon_longiceps,Tropeognathus_cf_robustus_SMNK_1133_PAL,((AMNH_22569,MN_6588_dash_V,(Tapejara_wellnhoferi,Vectidraco_daisymorrisae)),(Toolebuc_pterosaur,(Coloborhynchus_speilbergi,Dsungaripterus_weii))))));