(Sphenodon_punctatus,(Owenetta_kitchingorum,Anthodon_serrarius,(Odontochelys_semitestacea,(Proganochelys_quenstedti,((Proterochersis_robusta,(Kayentachelys_aprix,Condorchelys_antiqua,Indochelys_spatulata,(Heckerochelys_romani,Eileanchelys_waldmani,((Siamochelys_peninsularis,((Dorsetochelys_delairi,(((((Platychelys_oberndorferi,(Notoemys_tlaxiacoensis,Notoemys_laticentralis,Notoemys_oxfordiensis,Notoemys_zapatocaensis)),(((Phrynops_geoffroanus,((Prochelidella_cerrobarcinae,Yaminuechelys_maior),(Chelus_fimbriatus,(Chelodina_colliei,Chelodina_longicollis)))),(Elseya_dentata,Myuchelys_latisternum)),(Araripemys_barretoi,(Pelomedusa_subrufa,(Erymnochelys_madagascariensis,Podocnemis_expansa))))),((Solnhofia_parsonsi,(Thalassemys_moseri,Santanachelys_gaffneyi)),(Xinjiangchelys_juggarensis,((Hangaiemys_hoburensis,(Judithemys_sukhanovi,(Ordosemys_leios,(Dracochelys_bicuspis,Sinemys_lens)))),((Toxochelys_latiremys,((Caretta_caretta,Chelonia_mydas),(Mesodermochelys_undulatus,Dermochelys_coriacea))),(Platysternon_megacephalum,((Protochelydra_zangerli,(Macroclemys_schmidti,Macroclemys_temminckii,Chelydra_serpentina)),((Emys_orbicularis,(Mongolemys_elegans,Chrysemys_picta,Trachemys_scripta,Geoclemys_hamiltonii,Echmatemys_wyomingensis,(Stylemys_nebraskensis,(Gopherus_polyphemus,Eurotestudo_hermanni,(Chelonoidis_gringorum,Chelonoidis_chilensis))))),(Emarginachelys_cretacea,Hoplochelys_crassa,(Baptemys_wyomingensis,Dermatemys_mawii),(Xenochelys_formosa,Staurotypus_triporcatus,(Sternotherus_odoratus,Kinosternon_flavescens))),(Basilemys_variolosa,Yehguia_tatsuensis,Adocus_beatus,((Apalone_spinifera,Plastomenus_aff_thomassii,Pelodiscus_sinensis,Lissemys_punctata),(Anosteira_ornata,Carettochelys_insculpta))))))))))),(Portlandemys_macdowelli,Plesiochelys_etalloni)),(Neurankylus_eximius,(Trinitichelys_hiatti,(Plesiobaena_antiqua,(Boremys_pulchra,Baena_arenosa,Chisternon_undatum)))))),(Pleurosternon_bullockii,(Glyptops_plicatulus,Dinochelys_whitei)))),(Chubutemys_copelloi,(((Niolamia_argentina,Gaffneylania_auricularis,Warkalania_carinaminor,Meiolania_platyceps,Ninjemys_oweni),(Otwayemys_cunicularius,Kallokibotion_bajazidi)),(Mongolochelys_efremovi,Peligrochelys_walshae))))))),(Palaeochersis_talampayensis,Australochelys_africanus))))));