(Plesiochelyidae,(Meiolania,((Chelydroidea,(Testudinoidea,Trionychoidea)),((Toxochelys,(Osteopygis,(Ctenochelys,(Allopleuron,(Puppigerus,Chelonia_mydas))))),((Desmatochelys,Protostega_gigas),(Corsochelys,(Mesodermochelys,(Eosphargis,(Psephophorus,Dermochelys)))))))));