(Ancestor,((((Lissemys_punctata,(Cycloderma_aubryi,Cycloderma_frenatum)),(Gilmoremys_lancensis,(Plastomenus_thomasii,((Hutchemys_rememdium,Hutchemys_arctochelys),(Hutchemys_tetraneton,Hutchemys_sterea))))),(Cyclanorbis_elegans,Cyclanorbis_senegalensis)),((((Amyda_cartilagineus,(Pelochelys_bibroni,Chitra_indica)),(Trionyx_triunguis,(Rafetus_euphraticus,((Apalone_ferox,(Apalone_muticus,Apalone_spiniferus)),(Gobiapalone_breviplastra,Gobiapalone_orlovi))))),(Nilssonia_formosus,(Palea_steindachneri,(Pelodiscus_sinensis,Dogania_subplanus)))),(Aspideretes_gangeticus,Aspideretes_hurum))));