(Euparkeria,(Herrerasaurus,(Preondactylus,((Dimorphodon,Peteinosaurus),((Anurognathus,Batrachognathus,Dendrorhynchoides,Jeholopterus),(((Austriadactylus,Raeticodactylus),(Pterorhynchus,(Scaphognathus,Sordes),(Dorygnathus,(Cacibupteryx,Nesodactylus,Rhamphorhynchus)),(Allkaruen_braincase,((Darwinopterus,Wukongopterus),(Eosipterus,Gegepterus,Ctenochasma,Pterodaustro,(Pterodactylus,Cycnorhamphus),(Pterodactylus_longicol,Gnathosaurus),(((Nurhachius,(Istiodactylus,Haopterus)),((Ludodactylus,(Ornithocheirus,Anhanguera,Coloborhynchus)),(Boreopterus,(Pteranodon,(Nyctosaurus,Muzquizopteryx))))),((Germanodactylus_rhamph,((Germanodactylus_crist,Noripterus),(Dsungaripterus,Tatal_pterosaur))),(Lonchodectes,((Tapejara,Tupandactylus,Sinopterus,Huaxiapterus),((Shenzhoupterus,Chaoyangopterus),(Tupuxuara,Thalassodromeus),(Quetzalcoatlus,Zhejiangopterus,Azhdarcho))))))))))),(Campylognathoides,Eudimorphodon)))))));