(Archaeopteryx,(Jeholornis,((Sapeornis,((Protopteryx,(Orienantius,shenqiornis,(Eopengornis,Parapengornis),(Vescornis,(Junornis,Eoenantiornis,Qiliania,Bohaiornis,Parabohaiornis,Longusunguis,Zhouornis,Gretcheniao,(Eocathayornis,Gobipteryx,Iberomesornis,CUGB_P1202,(Concornis,Eoalulavis,Cathayornis),(Neuquenornis,Sulcavis))),((Elsornis,Longipteryx),(Shanweiniao,(Longirostravis,Rapaxavis))),(Pengornis,Fortunguavis)))),(Schizooura,Bellulornis,((Jianchangornis,(((Longicrusavis,(Apsaravis,Hongshanornis)),((Gansus_yumenensis,Gansus_zheni),(Ichthyornis,((Hesperornis,Parahesperornis),(Vegavis,(Anas,Gallus)))))),(Yanornis,Yixianornis))),(Archaeorhynchus,Patagopteryx))))),(Eoconfuciusornis,(Confuciusornis,Changchengornis,Vorona)))));