(Dipterus_valenciennesi,(Sagenodus_spp,(Beltanodus_ambilobensis,Gnathorhiza_spp,(Microceratodus_angolensis,(Neoceratodus_forsteri,((Ceratodus_sturii,(Ptychoceratodus_philippsi,(Arganodus_atlantis,Gosfordia_truncata,Protopterus_spp,Ptychoceratodus_serratus)),(Asiatoceratodus_sharovi,Paraceratodus_germanini)),(Ferganoceratodus_jurassicus,Ferganoceratodus_martini)))))));