(Homogalax_protapirinus,(Heptodon_calciculus,(Hesperaletes,(Dilophodon_minusculus,((Thuliadanta_mayri,(Helaletes_nanus,(Plesiocolopirus_hancocki,(Protapirus,((Paracolodon_fissus,Paracolodon_inceptus),(Colodon_stovalli,((Colodon_kayi,Colodon_woodi),(Colodon_cingulatus,(Colodon_occidentalis,Colodon_orientalis))))))))),(Heteraletes_lectanus,(Desmatotherium_intermedius,Desmatotherium_mongoliense)))))));