#NEXUS [File downloaded from graemetlloyd.com] BEGIN DATA; DIMENSIONS NTAX=47 NCHAR=424 ; FORMAT DATATYPE=STANDARD SYMBOLS=" 0 1 2 3 4 5 6 7" MISSING=? GAP=- ; MATRIX Simonsius ?????200?1?1?120?111020111001111011111100001?201?100111001210?1110?11011010001001?11?3????0??????0????01?00021002012?0210121121111100100103000?131220111000???0001021110111102100111100001111110112100??????????????1??122010?000000010000000021000200000?0110211(01)2(12)001110011???1??????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????22(01)1000----01020112101102 Cebuella 11010(12)0011012210?1010110(01)1200?111011121000110201111111100121021110?0101100220000111011101122121110101111020021000022?1200120010100000100102011?2212?110?10????00??020110000111233??11000?11?2000215010100102100000001100010(01)10000100200110001?100000022?0???0201102100?00??100?30223?(01)?(12)?020??0??200(12)21?(01)1(01)??2112202110020????2??120100(34)(12)(12)(12)0111?10?2100?201020100000??00110000??11?000112110001122?200110120011-01102210220020-0(01)200112- Callithrix (01)101000011012210?101011(01)01200?111(01)11121000110201111111100121021110?010110022000011101110122212111010122(12)020021000022?120002001010000010(01)102111?23(01)3?110?00????00??022110000110233??11000?11?20001140(01)01002210000000(01)1?00012010000100200110001?10000(01)022?0??00201002100?0(01)??100?(23)0(23)23?(01)?(12)?020??0??20(01)(12)2(01)?(01)0(01)??211?201110020????2??(12)20100(23)(12)2(12)(01)11111012000120102010???????????????????????????????????????????????-0110121032002200(01)2011211 Saguinus (01)1001000110122000101010011(12)00?111(01)11121000110201111011100121021110?0101100220100112101001210010000010222000021000022?100012011000000010(01)001011?13(01)2?(01)10?(01)0????00??01210(01)000110233??1(01)011?11?200110(45)110100211100010001100000100000(01)00210010001?0000000(12)2?0???0201001100?0(01)??20(01)010(12)1(01)1(01)1(012)1020010??20(01)(12)2(01)0(01)0(01)10211(12)(01)0111002(01)????2??12(01)1003(12)2(12)(01)11(012)?10?2100?201020100002?001110000?011?000102110001022?202111100010-00-02011331012001201222- Leontopithecus 0001100011012210?101010(01)00200?111111101000110201111111100121021110?0101100221001102111001120000000010212000021000022?1210(01)20110000000100102011?1304?110?10????00??0221000021102(23)(23)?010011?11?20001051(01)0100101100010001100011111000000210001001?(12)00000022?0???020101200??00??10(01)?(12)0(23)23?(12)?(12)?020??0????112(01)?(01)?1??2122101110020?0??200????003(12)2(12)(01)11(01)110121001201020100020?101110000?011?000101110001122?202110100010-00-022(01)022101300021111(01)(01) Lagonimico ?????000?101???????1??0(12)????????????1????0???20???11111???????1111???0????????01112211001120010000010012?00021000022?1000120110000010100001011?1????0110201???00?00221000001112222000012011120011141101002022000000011001110110001002100110012100000022?0???0001002(01)000000010??????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????-00-0121133112300121(01)2102 Callimico 1000010011012210?1010000(01)1100?111011121000110201111011100121021110?01?110122110010210100102000000001020(12)000021000022?1010220110000000100003011?1302?0110100???00?00221000001102333?2000001202000115110100102100000001??0102000000000210010000110000101200-001201002(01)0000000100?(23)020(01)?1?(12)?020??0??201(01)2(01)?(012)1(01)??210?1011100(02)(01)????200120100(23)(12)2(12)1(01)11?10?2000?201020100022?102100000?011?0001021000012120202120110011011100111331?2200021(01)1100 Cebus 1000(01)00011012200?1010(12)01(12)1000?021010121100201201110011100121021110?0101100222?(01)0112101001210000000010223100021000022?111(12)2201(12)0000000100102011?1303?0110(01)00???00?00211000001112332?000222110201010320010012000002101102?11220100001(01)21100000121100000000210?10111(01)(12)0000111121(01)?1021(01)12?0?020010??1(01)(01)(12)(23)(012)?(01)0(01)??(012)11(12)(12)01110011????200120100(34)(012)(23)(12)(01)1111001?0001?000201000220112101200101000002111000013011202110100001110-0100?112121102210222- Saimiri 111110001101220001010(12)0011000?111111121210201201111011100121021110?0101100222101112111001(12)(01)1000000010222200021000022?1210220220000010100000011?1303?0110(01)00???00?00221100001112(23)(23)10110001120200(12)114100100221000020011120011(12)010001002100010012(01)100000000100?00010220000010(01)10102020012102020010001(01)1(01)210(01)012021122021100201110100210000(34)12(123)01111001200012000201100121112101200101000121021100013011212121120011100-0111133(12)(02)2300(01)22011(12)2 Neosaimiri ?????????????????????????????????????????????????????????????????????????????10111???1001011000000010201200021000022?121022022000001010(01)000(01)11?1301?011?101???00?00221100001112(23)(23)?0110122121200(12)1141201??20(01)1000200110200120010001(01)021000100021100000(01)(01)00?0?0001002(01)000010010102010012102????????????????????????201110020???020???????????????????????????????????????????????????????????????????????????????10(01)11220011(01)(01)230112101112 Aotus 10010(01)0111012200?101010111100?0(12)1111121102210201111011100121021110?0101101211101112(12)01001220000000010221000021000022?121(01)220120000000100001011?1201?0110101???00?00211000001112333?10022211020001131001002220000200(01)1011112(01)0100000021000000022100000000200?000100(01)0000010(01)21(01)01(01)(123)101(01)1(012)10200100020(01)(012)2(01)0(01)1(01)20(012)11?10211001(12)1110100221000(23)(12)2(12)(01)11110012000120002011002211021000001011?001102110001322?202110110011200-02200332(12)20-00210212- Aotus_dindensis ??????01?????????????????????????????????????????????????????????????????????1011????1001111010000010011?01021000022?1010120110000010100002011?1311?011010????00?00211000001112333?100222111100011?????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????? Mohanamico ?????????????????????????????????????????????????????????????????????????????1011????1001??1?100000100?2?01021000022?1010120110000010100002011?1311?0110101???00??021100000111233??10022?11?100011?130?????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????? Callicebus 0001011011112220?101000111200?021(01)11121000210201111011100121021110?0101101221111102201001011010000010001000021000022?121(01)(12)20220000000100002011?1300?0110101???00?0021100000111(12)(23)(23)111102221101(01)0(01)113100100111000021011021111001000011210001000121000(01)00002000(01)0110020000001(01)1110(12)(01)213111(01)102001000201(01)2(01)000(01)20(12)11?10211001(01)1110100221000(23)12(12)01111001200012000201100220112110000101000011021100012222202110100001210-0001133(01)(01)220002001101 Pithecia 110110001101222011010(01)0111100?02(01)111111000200201110011100121021110?0101101222111112211001112020110000012101021000022?221(01)220220000000110003011?1301?0110101???00?01201000021112332?00022210020000032001002111000110110000111110000(01)121000000122100000000200?001110100001011210?30(12)001????02???0??200(12)(23)1?011??(01)111101110011????000120100322(12)0111?00?2000?200010110132010210110000000111111100001312?102110100011210-01110112222001210312- Cacajao 1111100011012221?101010121100?021011101100200201110011100121021110?01?1101222111112211001102120110000013201021000022?2110220220000000110003011?1303?0110101???00?02201000221102332?000222100200000320010022110001100101???1201010?1120100000112100000000200?0011100000000212(12)(01)?1(01)201?2?(012)?021??0????1(12)3(01)?(01)?(01)??(012)11?00211001(12)?????00?????0(34)12(12)011(01)?00?2000?20000010??????????????????????111100101212?102110100011(23)00-01111331120-22200332- Chiropotes 1?1??00011012220?101000(12)21100?0(12)1111101100200201111011100121021110?01?1101222111122211001102120110000013001021000022?2211220210000000110003011?1303?0110101???00?02201000221102332?000222100200000310010022100001100101???120101000120100000??2100000000200?0011100000010012(12)(01)?1120(01)?2?0?021??0??201(12)(23)(01)?(01)21??(012)11(12)001110001???????????00(34)(012)2(12)0111?00??000??0001011???????????????????????????????????????????????(23)00-01111331120-12200332- Nuciruptor ?????????????1???????????????????????????????????????????????????????????????1111????1001?12?201100?0001?00021000022?1?10221120000000100102011?1301?011010????00??020100002111233??00022?11?201000?200?????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????(34)???????????????????????????????????????????????????????????????????????????????????????????????? Proteropithecia ??????????????????????????????????????????????????????????????????????????????????????00???2120??00000?3?0002100??????2?0??????????????01?????????????1?1?????00??12?110000111233??10022?111200111?200?????????????1?11????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????? Alouatta 10000100(12)12120201101000221200?011(01)01001000210201110011100111021110?0101101220111122211011000000000010101200021000022?1210220(12)2000001?100000011?13(01)(12)?0111(12)0(12)???0000022(01)10000111(12)(23)(23)1021(01)112121(12)0002213(01)(01)100000100010001001(12)(012)00100000(012)(01)2100000(01)022200000000100?(01)11110(01)2000011(12)22(01)?102131(01)?02020??0????11(23)(01)?(01)?(01)??(01)222000110011????000120100(34)2(12)(23)0(01)1(12)?00?2000?200111111012?102100000?011?0011021100012021210120120011102001111330(01)10-21210222- Brachyteles 11011200(12)1212120?101000211200?011101011100200201110011100121021110?0111101?2?111122211001000000000010201200021000022?121022112101001?00000001101301?001?101???00000210110001122332?2110021220000123310100000101021011001110(01)10000000201000011221000000100?1?(01)21110000000011220?(12)022310?22021??1????(01)(12)(23)0?(01)?0??122??0011000(12)????????????042220(01)(01)1?00?2010?200121010012?002110100?01100011010100013220212120120011101001111330020-0(12)200222- Stirtonia ????????????????????????????????????????????????????????????????????????????????1????1???????0???00?0?0?????????????????????????0???????????????????011?2?????00??02?1100001112???020012?12?200011?301?????????????01001(12)(12)101000001120100001012200000000101?01010112000011122??????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????1020012113310(12)212121022(12)3 Ateles 110(01)1100111121200101000111100?0(12)1101011100200201110011100121021110?01(01)1101222111122211001200000000010122100021000022?121022011000001?100102011?130(12)?0000101???00?00201000001112232?000122111201011320010012100102101112000110000000021000000222(12)00000000101?00111000000001(01)22(01)?10(123)231(01)1(012)2021101????(01)(12)(23)(01)?(01)?(01)??(012)12200011000(01)?01010022?100(34)2(123)(123)011(01)100120101200121010032?002110100?01001011111000211121102111100011200-01111331020-00200222- Lagothrix 110(01)020011112110?101000211(12)00?021(01)01011100210201110011100121021110?0111101222111122211001210000000010222200021000022?121022012000001?100001011?1301?0110101???00?00201000001112332?1000221112000113200?00121101021011???00210000000021000000?22(12)00000000(01)00?001110000000011220?(23)02231(12)?02021???????1(12)3(01)?(01)?1??(012)222001110011????0??2201?04(01)(127)(12)01111001200012001210100220012110000101100011111000013222202120111011200-00111331020-00200332- Mazzonicebus ??????????????????0??????????????????????0????????????????????????????????????111????1001102120110000023?000210000?2?12?0?20?11?01000100102011??201?0????0?1??00??020111000?11233??01011?11?101100?2(01)0?????????????(01)11000021(01)0000022210101000?210(01)000110(01)0??00000(01)2(01)00?00??2????????????????????????????????????????1?????????????????????????????????????????????????????????????????112012001301121212112100010(01)10111133(01)(12)2(23)01121021(01)(23) Aegyptopithecus 0?00?0002101?020?001110121000?020211?0100(02)21?201?10011101121021110?01011001201001021110010?00100000101012000211?2022?1?1?02001010000010?103011?240120111101???010102110010000100001000000111111(12)111300?????????????0311?22?1?1000?21210001000012000110000?0010010(01)210001100121010(01)0011101010010???0?(12)2?0??0?????2?????????111011112110030131?12??0?20????0??????1?2??0?2?1?2?0?1?1120?102111001300121012212100011(01)101100111(12)020011101101 Apidium ?????00??1???1???101??????????1(12)??????1??011?201?101111?0?21021110????110?0??1001011110010?0000000010201200021002022?0001121001111101001101000?132(12)20111101???0002021110111101000010200000111111212100???????0??????1??222110100003100000000001200022001(01)01120211221002110111110(01)(12)0211101010010??211010010100(12)1?020211001(01)10101002101002021011(12)?????0????0?????????????????????????????????2???????????????????130-000001102030012102201 Catopithecus ?????000?1???1???101110121????1(12)0?????11?011?201?1???11001210???10?01011?12??100002?01?0???0010000010?012000211?201121?1?02(01)02010000010?00201101331201111011??000100100100011111110000000122200011301010000110002000310012?1??000?00200010010012000101100?0000011022000001011101(01)0001110101??????1210?00?000????2?????????11?01111211??????????????????????????????????????????????????????????????????????????00(01)102200111(12)020112111112 Proteopithecus ?????000?1?1?2?0?10?010121100?12021???11?011?201?1?0111001210?1110?010?1?12??100001101????????????????0?2???210022212101002101010000001210210101110101111001??000(01)012001000111111100000001222001113010???1???0?????0111?22?20?000000?01110?10101000001100?0?(01)001102200000001011011121110101?0????221010???10????11??1100??101010?1201??????????????????????????????????????????????????????????????????????????000-1220011110201122111(12)(23) Dolichocebus ??0?00?0?1?12200?10?02002??00??20???121?0010?2?1??1?1110012102111??0101101222?0??????1?0?0?0000??0010201200021000022?12101211201010001001000?1?1312?01111??1??00??021111000111022?111012?11?111110311010???1?0001000111011000100012220110100011200010000(01)0?002010020001011111?????????????????????????????????????????????1110100??????????????????????????????????????????????????????????????????????????????10(01)10111133(01)1220112101112 Branisella ?????????????1???????????????????????????0????????????????????????????????????001????10010(12)?000100010201100021000022?1210120110000000100003001?1113?0111101???000(01)021100000111111101101101121000113100?????????????01110110011000000201101000111000000000?0?(01)(02)111(01)2(01)000000021??????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????1(01)(01)1011113301(12)201121011(12)(23) Soriacebus ??????????????????????????????????????????????????????????????????????????????111????1001202120110000023?00021000022?1110120120101000100101011?2312?0110111???00?2021110000110122110001111112010003120?????????????0110122110000002221010100011101000110100?00101(01)2(01)000001021??????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????10(01)1011113311220112002123 Homunculus 110110002121?120?101020221000011121112110011?201?10011100121021110?01(01)11012211001(01)2111001001010110010001?00021000022?121(01)(12)21120?000001001020?1?1300?0111100???00?2022(01)(01)1000111(01)(23)(23)?011111?1(12)?200011220010011??0?0??01111001101000001221010000011100010000100000010(01)2(01)00100102100202011110100?01?????1121?0?1??2??2?????????????????????????????????????????????????????????????????????102012011312?212120120010(01)(01)0-011113322220112102101 Carlocebus ??????????????????????????????????????????????????????????????????????????????00?????10?1????????0?????1?0???10000211?2101201(12)01010?0?02102011?130??011?100???00?002?11000011102??111111?12?2100114200??????????????111?(01)1?0?0?00?(123)22101?100(01)?11000(01)0000100??0010(01)0(01)000000121??????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????10(01)1000113312220(01)12102101 Hylobates 1000000021012120?1010?01210000021111121000210201110011101121021111?0101100022100122111001110000000010?120000211?2222???1?0201101000??10?0030111242110111101???00010201001100010000100000011111101133?0100????0?????0312?22?1?0000?0021?00?100?22000000000-0?000100000?00?22222?3112311112??110?????1130?0?1??0223001110001?????112201?031301121?????0?????0??????????????1?1???1?1010?1011001102002100110101001200-0220000(01)202000220322- Miopithecus 1111100021012110?1010001110020021111121100200201110011101121021111?0101100022100122111001221000000010?212000211?2222???1?0201101000??10?0030111242010111101???00?00201001000022332?20000012200001133?0100????0?????0310?22?0?0000?00210000100?220000000020??001100000?01012222????2?022?????11???????????????11?200011002??10011101010030??1121?????0?????0????????????????????????????????????????????????????300-01111332200-00210322- Presbytis 1111100021012010?1010001210000021111121000210201110011101121021111?01011000221001221110011110000000100212000211?2222???1?0201101000??10?0030111231110111101???00010201001000022330?20000012200001123?0100????0?????0310?22?0??000?00210000100?220000000020??001100000?00022222?20223022????0111????1220?1?1??111100?110020?????110201?0302111210?11100????0101?????????????????????????????????????????????????200-??????????0-00200212- Tremacebus ??0??0??2111?100?10??1002?200?0200??12100111?20??11?11100121??111??010???1222?????????????????????????0??????10????????????????????????01???????????????????????????????????????????????????????????????????????????????11??????????????????121100???010100????1?12???????(01)20??????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????1???011113311220??210??12 Acrecebus ??????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????2?00000?0021??201?102000021??2???????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????210-0?0?1?321220002201102 Cebupithecia ??0??????????1?0?10??102??????????????1??0????????0011100121021110???001?1???1111????1001?021201100?0003?01021000022?1?1?22011000000010?102011?1300?0110?0????000??2010?00?1?123???00?2??11?20100?4100??0??????????010111110010000002100000002(12)00000001020??0001002000?102??0????????????????????????????????21??10?1100?1?0??100021?0?3????111?????????????2??????????????????????????????????????????????????200-01211331022001210220(23) Chilecebus ??????????????0??1?1????1??????????????1?0?(12)?2??????????????????10??????????0??????????????????????????1????????????????????????????????????????????????????????????????????????????????????????????????100220??2?011020110121000111?10101000?11000100000?000001002(01)000010011??????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????100-000--3322220102201101 Antillothrix 1101100011212120?101020221200001121111100021?201011?11101121021110?010110112?1101????2?????????0??????02?????00000??????????????????????????????????011010????????02?110000111133??011(12)(12)??1?201022320110?202000000011020(01)120110001002100010012(12)1000000100?0?00011(01)20000010(01)11?1100011210202021???100220000011????000110011?????????????????????????????????????????????????????????????????????????????????????000-001113310220112102112 Paralouatta ?00??1001??12021?101?202??1???02?1??1?1000?1?2???11?111001?????110?010???????111122211001020000000000000?0021100002?20200110110101010000100012?1313?0112111?????000201100001111332?11122212221002243211012021000200?1??(01)(12)(12)?211000100210001010(12)1100000001100?020112220000111121010111121111201????????20?????0???3001110011??????0???????????????????00??210????????????????????????????????????????????????????00110111133002202122022(01)(23) Xenothrix ??0????0?111?1?1?1010???0?????02?20??????0?1?????????????????????????????????110112112??1??????0?0????01?????1000022?0?1?22???001?01010?10000??1?00?111?01?????2??12?100011011000?200122?11?2110115200100(12)??????????1??011(01)0?1000?0021000000002000?0001020??0011100000021??2010100011(12)1111200????????3???0000??????????????????????????????????????????????????????????????????????????????????????????????????100-00?1?3??110-21210332- Panamacebus ?????????????????????????????????????????????????????????????????????????????????????1001?21000?001102?1?00021000022?1211?(12)0?10?0(01)010101000010???1????????????????????????????????????????????????????????????????????????????????????????????10000001(12)?10?-?????0?(01)00?01??????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????100-01100111(12)230002001102 Perupithecus ??????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????1-000(01)01??0-?-???1?1?(01)00?0???????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????-00-02?1?1?22030112121112 Canaanimico ??????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????11010001(12)?(01)0?-?????1?000?00??????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????100-0111133212(23)0112002223 ; END; BEGIN ASSUMPTIONS; OPTIONS DEFTYPE=unord PolyTcount=MINSTEPS ; TYPESET * UNTITLED = ord: 6 11-15 22 24-25 27 32 34 38 40 42-44 57 76-77 79 82-84 86 90-92 94 96-98 102-105 109 113-118 121-123 125-126 136 138-139 141-147 153 155-158 160 162-165 168 174-182 187-189 192-197 199 202-206 209 215-221 227-229 237-242 244-247 249 251 253 258-260 263-264 266-270 272 274-275 278 280-281 283 290-291 293-295 297-298 300 302-306 308-310 313-314 316 319 322-324 326 328-331 333-336 338 340 345 349 354-355 359-360 363-364 369-374 377 380-381 384 386-390 392-393 395 400-402 405-406 411-414 416-424, unord: 1-5 7-10 16-21 23 26 28-31 33 35-37 39 41 45-56 58-75 78 80-81 85 87-89 93 95 99-101 106-108 110-112 119-120 124 127-135 137 140 148-152 154 159 161 166-167 169-173 183-186 190-191 198 200-201 207-208 210-214 222-226 230-236 243 248 250 252 254-257 261-262 265 271 273 276-277 279 282 284-289 292 296 299 301 307 311-312 315 317-318 320-321 325 327 332 337 339 341-344 346-348 350-353 356-358 361-362 365-368 375-376 378-379 382-383 385 391 394 396-399 403-404 407-410 415; WTSET * UNTITLED = 12: 146-147, 15: 145 195 328, 20: 86 91 104 117 139 156-158 176-178 196-197 220 272 274 294 305-306 330-331 384 400-401 414 421-422 424, 30: 6 11 14-15 22 24-25 27 32 34 38 40 42-44 57 76-77 82-84 90 92 94 96-98 102-103 105 109 114-116 118 121-123 125-126 136 138 141-144 153 155 160 162-165 168 174-175 179-182 187-189 192-194 199 202-206 209 215-219 221 228-229 237-242 244-247 249 251 253 258-260 263-264 266-270 278 280-281 283 290-291 293 295 297-298 300 302 304 308-310 313-314 319 322-324 326 329 332-336 338 340 349 354-355 359-360 363-364 377 379 385-389 392-393 395 402 405-406 411-413 416-420 423, 60: 1-5 7-10 12-13 16-21 23 26 28-31 33 35-37 39 41 45-56 58-75 78-81 85 87-89 93 95 99-101 106-108 110-113 119-120 124 127-135 137 140 148-152 154 159 161 166-167 169-173 183-186 190-191 198 200-201 207-208 210-214 222-227 230-236 243 248 250 252 254-257 261-262 265 271 273 275-277 279 282 284-289 292 296 299 301 303 307 311-312 315-318 320-321 325 327 337 339 341-348 350-353 356-358 361-362 365-376 378 380-383 390-391 394 396-399 403-404 407-410 415; END;