(Biretia_megalopsis,(Biretia_piveteaui,Biretia_fayumensis,((Simonsius_grangeri,Parapithecus_fraasi),(Qatrania_wingi,Qatrania_basiodontos,(Apidium_bowni,(Apidium_moustafai,(Apidium_zuetina,Apidium_phiomense)))))));