(Manouria,((Gopherus_laticuneus,(Gopherus_agassizii,Gopherus_berlandieri,(Gopherus_brevisternus,Gopherus_pansus,Gopherus_vagus,(Gopherus_flavomarginatus,(Gopherus_canyonensis,Gopherus_polyphemus,(Gopherus_donlaloi,Gopherus_hexagonatus)))))),(Gopherus_mohavetus,(Gopherus_edae,Testudininae,(Hesperotestudo,Stylemys)))));