(Parahippus_leonensis,(M_gunteri,(M_primus,(((M_stylodontus,(M_carrizoensis,(Pliohippus,(Astrohippus,(Onohippidium,(Hippidion,(Equus,Dinohippus))))))),(Merychippus_isonesus,M_sejunctus)),(Protohippus,Calippus),(M_intermontanus,((M_coloradense,(Pseudhipparion,Neohipparion)),(M_insignis,(Hipparion,(M_goorisi,(Cormohipparion,Nannippus))))))))));