(Lambdotherium,(Eotitanops,(Lophiotherium,(Propalaeotherium_hassiacum,Propalaeotherium_messelense,(Palaeotherium,(Plagiolophus_annectens,Plagiolophus_sp_I)),(Propachynolophus_gaudryi,(Propachynolophus_maldani,(Hyracotherium_leporinum,(Anchilophus,(Pachynolophus,(H_sp_Rians,(H_tapirinum,(H_vasacciense,(H_vulpiceps,(H_aff_vulpiceps,(Cymbalophus,(Homogalax_sp_I,(Homogalax_protapirinus,(Isectolophus,(Kalakotia,(Breviodon,(Eoletes,((Rhodopagus,Teleolophus,(Chasmotherium,Protapirus,(Colodon_occidentalis,(Helaletes,((Triplopus,Heptodon),(Hyrachyus,(Eomoropus,(Lophiaspis_maurettei,(Paralophiodon_buchswillanum,Lophiodon))))))))),(Lophialetes,Schlosseria)))))))))))))))))))))));