(Manouria,(Gopherus_mohavetus,Gopherus_praecedens,(Gopherus_laticuneus,(Gopherus_agassizii,Gopherus_berlandieri,(Gopherus_brevisternus,Gopherus_pansus,Gopherus_vagus,(Gopherus_flavomarginatus,(Gopherus_canyonensis,Gopherus_polyphemus))))),(Gopherus_hexagonatus,(Gopherus_edae,(Stylemys,(Hesperotestudo,Testudininae))))));