Tapirus_terrestrisHyrachyus_eximiusTrigonias_osborniRonzotherium_filholiPlesiaceratherium_mirallesiProtaceratherium_minutumDiceratherium_armatumMenoceras_arikarenseKenyatherium_bishopiBugtirhinus_praecursorCaementodon_caucasicumCaementodon_oettingenaeHispanotherium_grimmiHispanotherium_beonenseHispanotherium_corcolenseHispanotherium_matritenseProcoelodonta_tekkayaiProcoelodonta_mongolienseHuaqingtherium_lintungenseIranotherium_morganiParelasmotherium_schansienseSinotherium_lagreliiElasmotherium_caucasicumElasmotherium_sibiricumOsteologyMaximum ParsimonyBased on reanalysis of the original matrix.Sanisidro_etal_2012a