AncestorCaptorhinidaeAraeoscelidiaClaudiosaurusYounginaRhynchosauriaTrilophosaurusProlacertiformesChoristoderaArchosauriformesKuehneosauridaeRhynchocephaliaSquamataCorosaurusCymatosaurusAugustasaurusPistosaurusPlesiosaurusSimosaurusGermanosaurusNothosaurusLariosaurusWumengosaurusSerpianoNeusticoAnaroDactyloKeichousaurusDianopachysaurusHovasaurusEusaurosphargisThalattosauriaYunguisaurusDiandongosaurusPsephodermaCyamodusPlacodusParaplacodusSinosaurosphargisWinterswijkSkullTestudinesHelveticosaurusOdontochelysHanosaurusIchthyopterygiaOsteologyMaximum ParsimonyBased on reanalysis of the original matrix.Neenan_etal_2013a