OutgroupPreondactylusDimorphodonPeteinosaurusAustriadactylusAnurognathusBatrachognathusDendrorhynchoidesJeholopterusCampylognathoidesEudimorphodonRaeticodactylusScaphognathusSordesPterorhynchusCacibupteryxDorygnathusAngustinaripterusSericipterusNesodactylusRhamphorhynchusQinglongopterusDarwinopterusPterodactylusOsteologyMaximum ParsimonyBased on reanalysis of the original matrix.Lu_etal_2012c