DromaeosauridaeArchaeopteryxJeholornisSapeornisConfuciusornis_sanctusChangchengornisEoconfuciusornisJinzhouornisConfuciusornis_duiBoluochiaConcornisElsornisEoalulavisCathayornisEocathayornisEoenantiornisGobipteryxIberomesornisLongipteryxLongirostravisNeuquenornisPengornisEopengornisProtopteryxRapaxavisShanweiniaoVescornisVoronaSchizoouraJianchangornisArchaeorhynchusSonglingornisLongicrusavisApsaravisHongshanornisYanornisPatagopteryxYixianornisPiscivoravisGansusIchthyornisHesperornisParahesperornsiEnaliornisBaptornis_advenusBaptornis_varneriVegavisAnasGallusFortunguavisQilianiaShenqiornisSulcavisBohaiornisParabohaiornisLongiungulavisZhouornisDunhuangiaIteravisPterygornisV20289OsteologyMaximum ParsimonyBased on reanalysis of the original matrix.Hu_et_OConnor_2017a