ProtungulatumArctocyonOrthaspidotheriumPleuraspidotheriumHyopsodusMeniscotheriumPhenacodusHeptodonHyracotheriumDiacodexisHomacodonRadinskyaTetraclaenodonEctocionHomogalaxHyracoideaProboscideaPhenacolophusLophocionOsteologyMaximum ParsimonyBased on reanalysis of the original matrix.Holbrook_2014ag