EuparkeriaHerrerasaurusPreondactylusDimorphodonPeteinosaurusAnurognathusBatrachognathusDendrorhynchoidesJeholopterusAustriadactylusCampylognathoidesEudimorphodonRaeticodactylusScaphognathusSordesPterorhynchusCacibupteryxDorygnathusNesodactylusRhamphorhynchusDarwinopterusWukongopterusPterodactylusCycnorhamphusEosipterusPterodactylus_longicolGnathosaurusGegepterusCtenochasmaPterodaustroIstiodactylusNurhachiusHaopterusOrnithocheirusAnhangueraColoborhynchusLudodactylusBoreopterusNyctosaurusMuzquizopteryxPteranodonGermanodactylus_cristGermanodactylus_rhamphDsungaripterusTatal_pterosaurNoripterusLonchodectesTapejaraTupandactylusSinopterusHuaxiapterusShenzhoupterusChaoyangopterusTupuxuaraThalassodromeusQuetzalcoatlusZhejiangopterusAzhdarchoAllkaruen_holotype_and_referredOsteologyMaximum ParsimonyBased on reanalysis of the original matrix.Codorniu_etal_2016ab