Tapirus_terrestrisHyrachyus_eximiusTrigonias_osborniSubhyracodon_occidentaleRonzotherium_filholiHispanotherium_beonenseBugtirhinus_praecursorCeratotherium_simumDiceros_bicornisDicerorhinus_sumatrensisRhinoceros_sondaicusRhinoceros_unicornisBrachypotherium_brachypusDiceratherium_armatumMenoceras_arikarensePlesiaceratherium_mirallesiProsantorhinus_douvilleiProtaceratherium_minutumTeleoceras_fossigerPleuroceros_pleurocerosDiaceratherium_aginenseAprotodon_fatehjangenseLartetotherium_sansanienseGaindatherium_browniAlicornops_simorrenseAlicornops_complanatumAlicornops_alfambrenseHoploaceratherium_tetradactylumChilotherium_anderssoniAcerorhinus_zernowiChilotherium_intermedium_ssAceratherium_incisivumOsteologyMaximum ParsimonyBased on reanalysis of the original matrix.Antoine_etal_2003a