(Eomoropus,(Moropus,(Butleria_rusingense,(Macrotheirum_brevirostris,((Anisodon_macedonicus,Macrotherium_macedonicum),(Macrotherium_grande,Styrie_Macrotherium_grande)),(Nestoritherium,(Chalicotherium_goldfussi_rhodanicum,(Chalicotherium_goldfussi,(Chalicotherium_wuduensis,mandibule_de_Titov_Veles))))))));