(Sphenodon_punctatus,(Simosaurus_gaillardoti,(Anthodon_serrarius,(Owenetta_kitchingorum,(Odontochelys_semitestacea,(Proganochelys_quenstedti,((Proterochersis_robusta,(Kayentachelys_aprix,Condorchelys_antiqua,Indochelys_spatulata,(Heckerochelys_romani,(Eileanchelys_waldmani,Chubutemys_copelloi,Mongolochelys_efremovi,Peligrochelys_walshae,Patagoniaemys_gasparinae,(Chengyunchelys,(Siamochelys_peninsularis,((Dorsetochelys_delairi,(((((Notoemys_laticentralis,(Platychelys_oberndorferi,Caribemys_oxfordiensis)),(((Phrynops_geoffroanus,((Prochelidella_cerrobarcinae,Yaminuechelys_maior),(Chelus_fimbriatus,(Chelodina_colliei,Chelodina_longicollis)))),(Elseya_dentata,Myuchelys_latisternum)),(Araripemys_barretoi,(Pelomedusa_subrufa,(Erymnochelys_madagascariensis,Podocnemis_expansa))))),(Xinjiangchelys_latimarginalis,(Hangaiemys_hoburensis,Judithemys_sukhanovi,(Solnhofia_parsonsi,(Thalassemys_moseri,Santanachelys_gaffneyi)),(Dracochelys_bicuspis,(Sinemys_lens,Ordosemys_leios)),((Toxochelys_latiremys,((Caretta_caretta,Chelonia_mydas),(Mesodermochelys_undulatus,Dermochelys_coriacea))),(Platysternon_megacephalum,((Protochelydra_zangerli,(Macroclemys_schmidti,Macroclemys_temminckii,Chelydra_serpentina)),((Emys_orbicularis,(Geoclemys_hamiltonii,Echmatemys_wyomingensis,((Mongolemys_elegans,(Stylemys_nebraskensis,(Gopherus_polyphemus,Eurotestudo_hermanni,(Chelonoidis_gringorum,Chelonoidis_chilensis)))),(Chrysemys_picta,Trachemys_scripta)))),((Emarginachelys_cretacea,((Baptemys_wyomingensis,Dermatemys_mawii),(Hoplochelys_crassa,(Xenochelys_formosa,Staurotypus_triporcatus,(Sternotherus_odoratus,Kinosternon_flavescens))))),((Shachemys_laosiana,(Adocus_beatus,(Basilemys_variolosa,Yehguia_tatsuensis))),(Anosteira_ornata,Carettochelys_insculpta,(Apalone_spinifera,Plastomenus_aff_dot__thomassii,Pelodiscus_sinensis,Lissemys_punctata))))))))))),(Portlandemys_macdowelli,Plesiochelys_etalloni)),(Neurankylus_eximius,(Trinitichelys_hiatti,(Plesiobaena_antiqua,(Boremys_pulchra,Baena_arenosa,Chisternon_undatum)))))),(Pleurosternon_bullockii,(Glyptops_plicatulus,Dinochelys_whitei))))),((Niolamia_argentina,Gaffneylania_auricularis,Warkalania_carinaminor,Meiolania_platyceps,Ninjemys_oweni),(Otwayemys_cunicularius,Kallokibotion_bajazidi)))))),(Palaeochersis_talampayensis,Australochelys_africanus))))))));