(S_krabiense,(((A_ulnifer,(A_thailandicus,(M_minimun,M_silistrensis))),(B_fraasi,((B_aequatorialis,B_onoideus),((A_americanus,B_velaunus),(E_armatus,(E_crispus,(E_borbonicus,(A_acridens,(A_acrid_Toledo_Bend,A_meridionalis)),(A_zelteni,((S_palaeindicus,H_blandfordi),((M_pusillus,(M_dissimilis,M_medioximus)),(L_anisae,L_petrocchii))))))))))),(H_occidentalis,(A_chaimanei,A_pangan,(A_magnum,A_mosvialense)))));