(Sphenodon_punctatus,(Owenetta_kitchingorum,Anthodon_serrarius,(Odontochelys_semitestacea,(Proganochelys_quenstedti,(Proterochersis_robusta,Palaeochersis_talampayensis,Australochelys_africanus,Kayentachelys_aprix,Condorchelys_antiqua,Heckerochelys_romani,Eileanchelys_waldmani,Indochelys_spatulata,Niolamia_argentina,Ninjemys_oweni,Warkalania_carinaminor,Meiolania_platyceps,Chubutemys_copelloi,Peligrochelys,Patagoniaemys_gasparinae,Spoochelys_ormondea,((((Glyptops_plicatulus,((Thalassemys_moseri,((Solnhofia_parsonsi,(Otwayemys_cunicularius,((((Emarginachelys_cretacea,(Emys_orbicularis,Shachemys_laosiana,(Geoclemys_hamiltonii,Echmatemys_wyomingensis,(Mongolemys_elegans,((Notoemys_laticentralis,(Platychelys_oberndorferi,Caribemys_oxfordiensis)),(((Phrynops_geoffroanus,(Prochelidella_cerrobarcinae,Yaminuechelys_maior),(Chelus_fimbriatus,(Chelodina_colliei,Chelodina_longicollis))),(Elseya_dentata,Myuchelys_latisternum)),(Araripemys_barretoi,(Pelomedusa_subrufa,(Erymnochelys_madagascariensis,Podocnemis_expansa)))))),(Chrysemys_picta,Trachemys_scripta,(Stylemys_nebraskensis,(Eurotestudo_hermanni,(Gopherus_polyphemus,(Chelonoidis_gringorum,Chelonoidis_chilensis)))))),((Baptemys_wyomingensis,Dermatemys_mawii),(Hoplochelys_crassa,(Xenochelys_formosa,Staurotypus_triporcatus,(Sternotherus_odoratus,Kinosternon_flavescens)))),(Basilemys_variolosa,Adocus_beatus,(Yehguia_tatsuensis,Basilochelys_macrobios)),(Anosteira_ornata,Carettochelys_insculpta,(Apalone_spinifera,Plastomenus_aff_thomassii,Pelodiscus_sinensis,Lissemys_punctata)))),(Macroclemys_schmidti,Macroclemys_temminckii,Protochelydra_zangerli,Chelydra_serpentina,Platysternon_megacephalum)),(Toxochelys_latiremys,((Caretta_caretta,Chelonia_mydas),(Mesodermochelys_undulatus,Dermochelys_coriacea)))),(Hangaiemys_hoburensis,Changmachelys_bohlini,Judithemys_sukhanovi,((Siamochelys_peninsularis,(Xinjiangchelys_wusu,Annemys_levensis,Annemys_latiens,Xinjiangchelys_radiplicatoides,Xinjiangchelys_junggarensis)),(Xiaochelys_ningchengensis,((Manchurochelys_manchoukuoensis,(Dracochelys_bicuspis,(Sinemys_brevispinus,Sinemys_lens,Sinemys_gamera))),(Ordosemys_leios,Liaochelys_jianchangensis)))))))),(Portlandemys_macdowelli,Plesiochelys_etalloni))),(Neurankylus_eximius,((Trinitichelys_hiatti,Plesiobaena_antiqua),(Boremys_pulchra,Baena_arenosa,Chisternon_undatum))))),((Dorsetochelys_delairi,Kallokibotion_bajazidi),(Pleurosternon_bullockii,Dorsetochelys_typocardium))),(Helochelydra_nopcsai,Naomichelys_speciosa)),(Sichuanchelys_chowi,(Sichuanchelys_palatodentata,(Mongolochelys_efremovi,Laurasichersis_relicta)))))))));