(Palaeosyops_robustus,(Telmatherium_validus,Microtitan_mongoliensis,(Epimanteoceras_formosus,Dolichorhinoides_angustidens),(Protitian_minor,Protitian_grangeri),(Gnathotitan_berkeyi,((Rhinotitan_kaiseni,Rhinotitan_mongoliensis),(Parabrontops_gobiensis,(Akautitian_hippopotamopus,((Metatitan_relictus,Metatitan_primus,Brachydoastematherium_transylvanicum),(Metatitan_progressus,Embolotherium_grangeri,Embolotherium_andrewsi))))))));