(Dimorphodon,(Rhamphorhynchus,((Cycnorhamphus,(Pterodactylus,Eosipterus,Ctenochasma,Pterodaustro)),(Haopterus,(Istiodactylus,Liaoxipterus),(Boreopterus,(Ornithocheirus,Coloborhynchus),(Nyctosaurus,Pteranodon))),(Germanodactylus,Dsungaripterus,Tatal_pterosaur,Noripterus),((Sinopterus,(Tapejara,Huaxiapterus)),(Shenzhoupterus,Chaoyangopterus,Jidapterus,Eopteranodon,Eoazhdarcho),(Tupuxuara,Thalassodromeus),(Quetzalcoatlus,Zhejiangopterus,Azhdarcho)))));