(Hypothetical_ancestor,(Proganochelys_quenstedti,(Proterochersis_robusta,Palaeochersis_talampayensis,Australochelys_africanus,(Kayentachelys_aprix,(Kallokibotion_bajazidi,(Meiolania_platyceps,Mongolochelys_efremovi),(Dorsetochelys_delairi,(Portlandemys_mcdowelli,Plesiochelys_solodurensis,(Xinjiangchelys_latimarginalis,(Hangaiemys_hoburensis,((((Platysternon_megacephalum,(Mongolemys_elegans,Gopherus_polyphemus,Chrysemys_picta,Geoclemys_hamiltonii,Peltochelys_durlstonensis,Sandownia_harrisi,((Platychelys_oberndorferi,Caribemys_oxfordiensis,Notoemys_laticentralis),(Elseya_dentata,(Chelodina_oblonga,Phrynops_geoffroanus)),(Pelomedusa_subrufa,(Erymnochelys_madagascariensis,Podocnemis_expansa))),(Emarginachelys_cretacea,((Baptemys_wyomingensis,Dermatemys_mawii),(Hoplochelys_crassa,(Staurotypus_triporcatus,(Sternotherus_odoratus,Kinosternon_flavescens))))),(Adocus_beatus,(Zangerlia_neimongolensis,Basilemys_variolosa)),((Apalone_spinifera,Lissemys_punctata),(Anosteira_ornata,Carettochelys_insculpta)))),(Protochelydra_zangerli,(Macroclemys_temminckii,Chelydra_serpentina))),(Toxochelys_latiremis,((Caretta_caretta,Chelonia_mydas),(Mesodermochelys_undulatus,Dermochelys_coriacea)))),(Judithemys_sukhanovi,(Dracochelys_bicuspis,(Sinemys_lens,Ordosemys_leios)))))),(Solnhofia_parsonsi,(Thalassemys_moseri,Santanachelys_gaffneyi))),((Pleurosternon_bullockii,(Glyptops_plicatulus,Dinochelys_whitei)),(Neurankylus_eximius,(Trinitichelys_hiatti,(Plesiobaena_antiqua,(Boremys_pulchra,(Baena_arenosa,Chisternon_undatum))))))))))));