(outgroup,(Macaca_sylvanus,Macaca_nemestrina,Papio,Theropithecus,Lophocebus,Allenopithecus,Miopithecus,Cercopithecus,Procynocephalus,Paradolichopithecus,Dinopithecus,Parapapio,Gorgopithecus,(Mandrillus,Cercocebus)));