(Cardiolophus_radinskyi,((Hallensia_matthesi,Hyracotherium_leporinum,Propachynolophus_gaudryi,(Propachynolophus_levei,Propachynolophus_maldani)),(Pliolophus_vulpiceps,Xenicohippus_osborni),(Hyracotherium_aff_cuniculus,(Pachynolophus_hookeri,Cymbalophus_cuniculus)),(Pachynolophus_eulaliensis_sp_nov,(Pachynolophus_cesserasicus,((Pachynolophus_duvali,(Pachynolophus_bretovensis,(Pachynolophus_garimondi,(Pachynolophus_lavocati,(Pachynolophus_zambranensis,(Paranchilophus_remyi,(Anchilophus_dumasi,((Eurohippus_parvulus,Lophiotherium_cervulum),(Palaeotherium_magnum,Plagiolophus_minor))))))))),(Pachynolophus_livinierensis,Pachynolophus_boixedatensis))))));