(Euparkeria_capensis,(Ornithosuchus_longidens,(Herrerasaurus_ischigualastensis,(Scleromochlus_taylori,(((Peteinosaurus_zambellii,((Eudimorphodon_cromptonellus,(Eudimorphodon_ranzii,Eudimorphodon_rosenfeldi)),(Caviramus_schesaplanensis,Raeticodactylus_filisurensis))),(Austriadactylus_cristatus,Preondactylus_buffarinii)),((Rhamphinion_jenkinsi,(Parapsicephalus_purdoni,(Dimorphodon_macronyx,Allkaruen_koi))),((Campylognathoides_liasicus,Campylognathoides_zitteli),((Dorygnathus_banthensis,((Fenghuangopterus_lii,(Qinglongopterus_guoi,(Harpactognathus_gentryii,(Sericipterus_wucaiwanensis,Angustinaripterus_longicephalus)))),(Rhamphorhynchus_muensteri,(Cacibupteryx_caribensis,Nesodactylus_hesperius)))),(Scaphognathus_crassirostris,(Orientognathus_chaoyangensis,(Sordes_pilosus,(((Kungpengopterus_sinensis,(Douzhanopterus_zhengi,Wukongopterus_lii)),(Pterorhynchus_wellnhoferi,((Darwinopterus_linglongtaensis,(Darwinopterus_modularis,Darwinopterus_rubostodens)),(Changchengopterus_pani,(Kryptodrakon_progenitor,(Painten_pterodactyloid,(((Pterodactylus_antiquus,((Kepodactylus_insperatus,(Elanodactylus_prolatus,(Ardeadactylus_longicollum,(Liaodactylus_primus,((Huanhepterus_quingyangensis,(Plataleorhynchus_streptophorodon,(Gnathosaurus_subulatus,(Gnathosaurus_macrurus,Lacusovagus_magnificens)))),(Moganopterus_zhuiana,Feilongus_youngi)))))),(Ctenochasma_elegans,(Pterodaustro_guinazui,(Beipiaopterus_chenianus,Gegepterus_changae))))),((Cycnorhamphus_suevicus,(Aurorazhdarcho_primordius,Normannognathus_wellnhoferi)),(Cuspicephalus_scarfi,(Germanodactylus_cristatus,Germanodactylus_rhamphastinus)))),(Haopterus_gracilis,(((((Aetodactylus_halli,(Cimoliopterus_cuvieri,Cimoliopterus_dunni)),(((Liaoningopterus_gui,(Anhanguera_piscator,(Anhanguera_blittersdorffi,(Tropeognathus_mesembrinus,(Ornithocheirus_simus,(Siroccopteryx_moroccensis,(Coloborhynchus_clavirostris,Coloborhynchus_wadleighi))))))),(Brasileodactylus_araripensis,Cearadactylus_atrox)),(Ludodactylus_sibbicki,Guidraco_venator))),(Nurhachius_ignaciobritoi,(Liaoxipterus_brachyognathus,(Istiodactylus_sinensis,Istiodactylus_latidens)))),(((Boreopterus_cuiae,Zhenyuanopterus_longiristris),(Lonchodectes_compressirostris,Lonchodraco_giganteus)),(Hongshanopterus_lacustris,((Ornithostoma_sedgwicki,(Pteranodon_longiceps,Pteranodon_sternbergi)),(Alamodactylus_byrdi,(Cretornis_hlavaci,(Muzquizopteryx_coahuilensis,(Nyctosaurus_lamegoi,(Nyctosaurus_gracilis,Nyctosaurus_nanus))))))))),((Bakonydraco_galaczi,(Bennettazhia_oregonensis,(Sinopterus_dongi,(Huaxiapterus_jii,((Eopteranodon_lii,((Tupandactylus_navigans,Tupandactylus_imperator),(Europejara_olcadesorum,(Caiuajara_dobruskii,(Tapejara_wellnhoferi,Vectidraco_daisymorrisae))))),(Huaxiapterus_benxiensis,Huaxiapterus_corallatus)))))),((((Dsungaripterus_weii,Domeykodactylus_ceciliae),(Noripterus_parvus,Noripterus_complicidens)),(Thalassodromeus_sethi,(Tupuxuara_longicristatus,Tupuxuara_leonardii))),((Eoazhdarcho_liaoxiensis,(Chaoyangopterus_zhangi,(Jidapterus_edentus,(LPM_L112113,(LPM_N081607,Shenzhoupterus_chaoyangensis))))),(Radiodactylus_langstoni,(Azhdarcho_lancicollis,(Zhejiangopterus_linhaiensis,(Arambourgiania_philadelphiae,Quetzalcoatlus)))))))))))))))),(Dendrorhynchoides_mutoudengensis,(Anurognathus_ammoni,(Batrachognathus_volans,(Jeholopterus_ningchengensis,Dendrorhynchoides_curvidentatus))))))))))))))));