(Outgroup,(Preondactylus,((Dimorphodon,Peteinosaurus),((Batrachognathus,Dendrorhynchoides,(Anurognathus,holopterus)),(((Austriadactylus,Raeticodactylus),(((Pterorhynchus,(Scaphognathus,Sordes)),(Darwinopterus,((Pterodactylus,(Cycnorhamphus,(Eosipterus,Gegepterus,Ctenochasma,(Pterodaustro,(Pterodactylus_longicolum,Gnathosaurus,Cearadactylus))))),(((Boreopterus,((Haopterus,(Istiodactylus,Nurhachius)),(Ludodactylus,Cearadactylus_atrox,(Ornithocheirus,Anhanguera,Coloborhynchus)))),(Pteranodon,(Nyctosaurus,Muzquizopteryx))),((Germanodactylus_rhamphastinus,((Germanodactylus_cristatus,Noripterus),(Dsungaripterus,Tatal_pterosaur))),(((Lonchodectes,(Shenzhoupterus,Chaoyangopterus),(Tupuxuara,Thalassodromeus)),(Quetzalcoatlus,Zhejiangopterus,Azhdarcho)),(Sinopterus,(Huaxiapterus,(Tapejara,Tupandactylus))))))))),(Cacibupteryx,(Dorygnathus,(Nesodactylus,Rhamphorhynchus))))),(Campylognathoides,Eudimorphodon))))));