(Sphenodon_punctatus,(Simosaurus_gaillardoti,(Anthodon_serrarius,(Owenetta_kitchingorum,(Odontochelys_semitestacea,(Proganochelys_quenstedti,((Proterochersis_robusta,(Indochelys_spatulata_2,(Kayentachelys_aprix,(Condorchelys_antiqua,(Heckerochelys_romani,(Eileanchelys_waldmani,((Chengyunchelys,(Siamochelys_peninsularis,((Dorsetochelys_delairi,(((((Notoemys_laticentralis,(Platychelys_oberndorferi,Caribemys_oxfordiensis)),(((Phrynops_geoffroanus,((Prochelidella_cerrobarcinae,Yaminuechelys_maior),(Chelus_fimbriatus,(Chelodina_colliei,Chelodina_longicollis)))),(Elseya_dentata,Myuchelys_latisternum)),(Araripemys_barretoi,(Pelomedusa_subrufa,(Erymnochelys_madagascariensis,Podocnemis_expansa))))),(Xinjiangchelys_latimarginalis,((Solnhofia_parsonsi,(Thalassemys_moseri,Santanachelys_gaffneyi)),((Hangaiemys_hoburensis,(Judithemys_sukhanovi,(Dracochelys_bicuspis,(Sinemys_lens,Ordosemys_leios)))),((Toxochelys_latiremys,((Caretta_caretta,Chelonia_mydas),(Mesodermochelys_undulatus,Dermochelys_coriacea))),(Platysternon_megacephalum,((Protochelydra_zangerli,(Macroclemys_temminckii,(Macroclemys_schmidti,Chelydra_serpentina))),((Emys_orbicularis_2,(Echmatemys_wyomingensis,(Geoclemys_hamiltonii,((Mongolemys_elegans,(Stylemys_nebraskensis,((Gopherus_polyphemus,Eurotestudo_hermanni_2),(Chelonoidis_gringorum,Chelonoidis_chilensis)))),(Chrysemys_picta,Trachemys_scripta))))),((Emarginachelys_cretacea,((Baptemys_wyomingensis,Dermatemys_mawii),(Hoplochelys_crassa,((Xenochelys_formosa,Staurotypus_triporcatus),(Sternotherus_odoratus,Kinosternon_flavescens))))),((Shachemys_laosiana_2,(Adocus_beatus,(Basilemys_variolosa,Yehguia_tatsuensis))),(Carettochelys_insculpta,(Anosteira_ornata,(Lissemys_punctata,(Plastomenus_aff_thomassii,(Apalone_spinifera,Pelodiscus_sinensis))))))))))))))),(Portlandemys_macdowelli,Plesiochelys_etalloni)),(Neurankylus_eximius,(Trinitichelys_hiatti,(Plesiobaena_antiqua,(Boremys_pulchra,(Baena_arenosa,Chisternon_undatum))))))),(Pleurosternon_bullockii,(Glyptops_plicatulus,Dinochelys_whitei))))),(Chubutemys_copelloi,(Mongolochelys_efremovi,((Patagoniaemys_gasparinae,(Otwayemys_cunicularius,Kallokibotion_bajazidi)),(Niolamia_argentina_et_Crossochelys_corniger,(Ninjemys_oweni,(Warkalania_carinaminor,Meiolania_platyceps))))))))))))),(Palaeochersis_talampayensis,Australochelys_africanus))))))));