(Conchopoma,(Aphelodus,Archaeoceratodus,Metaceratodus,Paraceratodus,(Megapleuron,(Gnathorhiza,(((Arganodus,Ferganoceratodus),(Asiatoceratodus,Gosfordia)),((Beltanodus,(Namatozodia,(Ariguna,Ceratodus),(Microceratodus,Ptychoceratodus))),(Neoceratodus,(Mioceratodus,(Lepidosiren,Protopterus)))))))));